Old Franks Fishpasses 1999 Monitoring Update
2-4-2000
Prince of Wales Island, Southeast Alaska
John Hannon, Craig Ranger District
This report summarizes the results of monitoring the Old Franks Fishpass Project during 1999 and makes comparisons with previous years. Monitoring efforts during 1999 focused on determining coho salmon escapement and colonization in the Old Franks system.
The 1999 monitoring activities included the following:
Two fishpasses were constructed over waterfalls on lower Old Franks Creek on Prince of Wales Island (Figure 1) in 1992. Coho salmon (Oncorhynchus kisutch) and pink salmon (O. gorbuscha) negotiated the fishpasses during the 1992 run. Coho moved into the upper reaches of the accessible habitat during that year while pink salmon did not make it to the upper fishpass. Pink salmon numbers have since increased and some pink salmon now make it through the upper fishpass. Chum salmon (O. Keta) have also begun to colonize the habitat between the fishpasses.
Data Available
The Alaska Department of Fish and Game began fisheries investigations in the Old Franks Watershed in 1978. These were augmented with additional fisheries data collected in 1989 (Zadina and Haddix 1993). The Forestry Sciences Lab collected baseline population and habitat data on the system in 1990 to later evaluate the response of the system to the introduction of anadromous salmonids (Bryant 1991). The Craig Ranger District of the US Forest Service began to monitor population and habitat conditions in the system in 1991 and has continued through 1999 (Hannon 1994, 1995, 1996, 1997, and 1998). The Ketchikan Area conducted a watershed analysis relating to timber harvest in the upper watershed in 1994 (USDA 1994).
Stocking History
Sockeye salmon (O. nerka) fingerlings were stocked in 1992 and coho fingerlings were stocked in 1994, 1995, and 1996 (Table 1). The coho bioenhancement program was a three year project conducted by the Alaska Department of Fish and Game (Denton 1997). Coho egg takes were conducted above Salmon Lake, in McGilvery Creek, in the Karta system. Karta is eleven miles north of Old Franks, in Kasaan Bay on the east side of Prince of Wales Island (Figure 1). Egg takes were conducted in late October and November. The fry were stocked into the Old Franks system the following summer.
Sockeye and king salmon were both stocked into the Old Franks system in the past, in the 1950's and 1960's. No sockeye or king salmon runs developed, but there is a kokanee population in the Old Franks Lake system which may have been established from the sockeye stocking. Presently no other stocking is planned. Resident salmonids present above the barrier waterfalls in the system prior to fishpass construction and stocking were cutthroat trout (O. clarki), rainbow trout (O. mykiss), and Dolly Varden char (Salvelinus malma).
Table 1. Stocking History of the Old Franks system.
| Date | Species | Age | Number | Location Planted | Source |
| 1952 | Sockeye | Green Eggs | 60,000 | Unknown | Buschmann Creek |
| 1952 | Sockeye | Eyed Eggs | 35,000 | Unknown | Buschmann Creek |
| 1953 | Sockeye | Green Eggs | 165,550 | Unknown | Buschmann Creek |
| 1962 | King Salmon | Fry | 33,750 | Lakes | Soos Cr., WA |
| 1963 | King Salmon | Fry | 46,223 | Lakes | Karta (Beaver Falls reared) |
| 5-14-92 | Sockeye | Fry | 227,200 | Lakes | Karta (Klawock H reared) |
| 8-16-94 | Coho | Fry | 96,632 | Upper Lake | Karta (Klawock H reared) |
| 7-21-95 | Coho | Fry | 78,719 | Bridge | Karta (Klawock H reared) |
| 7-26-95 | Coho | Fry | 139,405* | O.F. Lake | Karta (Klawock H reared) |
| 6-25-96 | Coho | Fry | 143,416 | O.F. Lake | Karta (Beaver Falls reared) |
34,735 cohos were coded wire tagged in 1994, 32,728 were tagged in 1995, and 44,431 were tagged in 1996.
*9,223 cohos with the same tag code were accidentally released into the Klawock River in 1995.
Methods
Site Description
The Old Franks watershed on Prince of Wales Island is 32 km east of Craig (Figure 1). It covers 65 km2 and flows into Polk Inlet (Figure 2). The anadromous lakes in the system cover over 300 ha and there are over 19 km of anadromous streams made accessible by the fishpasses.
Escapement Counts
1999 escapement
We captured coho in lower Old Franks Creek, between saltwater and Lake Mary on six days between July 22 and September 2. A monofilament gillnet 16 m long and 2.4 m deep with 44 mm square mesh was drifted through pools with holding coho. A snorkeler identified pools for netting and aided in enticing fish into the net. Healthy coho were tagged below the dorsal fin with a 30 cm yellow spaghetti tag. The recapture event was conducted on October 4 by snorkeling upper Old Franks Creek and counting marked and unmarked coho. A Peterson estimate was used to estimate the population.
1993-1998 escapements
A Smith Root Model 1100 Fish Counter was installed in a horizontal flume at the upper fishpass and was used to count returning adult salmon between 1993 and 1998. A picket fish trap was installed in the exit pool of the fishpass, above the fish counter. The trap helped to validate counts made on the fish counter. Adult salmon captured in the picket trap were measured (mid-eye to fork length), examined for adipose clips, and tagged with a numbered 30 cm yellow spaghetti tag. The tag allowed us to identify fish upstream that passed through the fishpass and to identify fish that went back down over the falls. We took scales from coho and sockeye that were captured in the picket trap. A small number (7-16) of the adipose-clipped coho were killed each year at the fishpass to recover the coded-wire tags. The tags identify which stocked group the fish is from. We also conducted periodic snorkel counts from Lake Mary down to saltwater to monitor the upstream migration of the salmon. A Peterson mark-recapture estimate (Chapman modification) was used to estimate escapement through both fishpasses. Confidence intervals were calculated using Appendix II of Ricker (1975). The recapture events were conducted by snorkeling upper Old Franks Creek and walking smaller tributary streams, primarily Trail Creek and Toque Creek.
Neff Creek and New Pipe Creek Fish Survey
Neff Creek and New Pipe Creek both flow into upper Old Franks Creek on the left bank (looking downstream) at 1,675 and 3,564 meters up from the lake respectively. These tributaries were established as coho density monitoring sites. Reaches were marked with orange tags and surveyed using the tier 3 stream survey (USDA Forest Service 1999). The marked reaches were trapped using minnow traps baited with salmon eggs. Three pass depletion estimates were used to estimate fish populations. Three trapping sessions were used with each trapping session lasting 90 minutes. Captured fish were measured and removed from the stream until the end of the third trapping session. Program CAPTURE (White et al. 1982) was used to calculate the population estimate and confidence intervals. Insufficient numbers of cutthroat or dolly varden were caught to conduct population estimates on those species.
Stream Population and Habitat Surveys
Habitat surveys were conducted in 1990, 1991, 1992, 1994,
and 1999 on upper Old Franks Creek from the upper lake to the top of anadromous
fish habitat. Fish densities (all ages and species) were determined annually
since 1990 by snorkeling established habitat units. New snorkel units were
marked periodically to adjust for natural changes in habitat units. The
1999 stream survey used the latest tier 3 stream survey protocol (USDA
Forest Service 1999). Forty-eight newly established snorkel sites were
marked with orange plastic tags on the upstream and downstream ends.
Results
Coho salmon escapement in Old Franks Creek
We trapped coho in lower Old Franks Creek on six occasions between July 22 and September 2. We marked 247 coho in lower Old Franks Creek, between saltwater and Lake Mary (Table 2). After September 2, pink salmon abundance prevented successful capture of coho. Figure 3 shows coho run timing since 1995. The 1999 timing appears earlier than in past years but the coho in 1999 were captured lower in the system than in past years. The coho timing in years prior to 1999 was based on captures in the exit pool of the upper fishpass. Coho routinely hold in the lower reaches of Old Franks Creek (lower 3 kilometers) in July and August before passing the upper fishpass.
We snorkeled upper Old Franks Creek on October 4 to count marked and unmarked coho for the recapture event. We counted 277 coho and seven of them were tagged. The escapement estimate is 8,582 (95% confidence interval = 4,458 – 18,069). This compares to estimates of 4,991 in 1998, 4,488 in 1997, 6,664 in 1996, 2,222 in 1995 and 250 in 1993 and 1994 (Figure 4).
The 1999 coho escapement estimate was the highest since the fishpass was installed. It consisted mostly of coho spawned naturally in the Old Franks system. Three percent of the coho from the 1995 egg take spent two years in the Old Franks system before out-migrating. Those fish returned in 1999. Most spent just one year in freshwater and returned in 1998. The remainder of returning coho in 1999 were wild fish spawned in Old Franks.
Table 2. Number of healthy coho tagged by date.
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Figure 3. Old Franks coho run timing.
Figure 4. Old Franks coho escapement estimates, 1993 - 1999. Error bars are 95% confidence intervals.
The Alaska Department of Fish and Game collected nine Old Franks tagged coho from the common property fishery in 1999. The estimated bioenhancement contribution, based on the tagging ratio and number of tagged fish collected, to the fisheries was 70 coho in 1999, 2,653 coho in 1998, 1,807 coho in 1997 and 1,877 coho in 1996 (Figure 5). Most coho in the commercial fishery were landed between July 15 and the end of August each year. Five sport caught coho were collected in 1998, three from Sitka and one each from Craig and Ketchikan. The estimated sport contribution in 1998 was 65 coho. Two sport caught coho were collected in 1997 at Sitka. Two sport caught coho were collected in 1996, one each in Sitka and Ketchikan. No sport contribution was estimated in 1996 or 1997.
Figure 5. Old Franks coho bioenhancement contribution to fisheries by date.
The majority of the bioenhanced coho completed their life cycle in three years, spending only one year in freshwater before outmigrating as smolts (Table 3).
Table 3. Percentage makeup of Old Franks coho bioenhancement returns by brood year source, based on coded wire tag returns.
| Brood Year | 1996 return | 1997 return | 1998 return | 1999 return |
| 1993 | 100% | 1% | ||
| 1994 | 99% | 8% | ||
| 1995 | 92% | 100% |
Table 4 shows the number of coho captured in the commercial fisheries by year harvested and year spawned (brood year).
Table 4. Coho bioenhancement contribution to commercial
fisheries (number of fish) by brood year (horizontal axis) and return year
(vertical axis).
Year Spawned (brood year)
| 1993 | 1994 | 1995 | Total | |
| 1996 | 1,877 | 1,877 | ||
| 1997 | 29 | 1,778 | 1,807 | |
| 1998 | 206 | 2,447 | 2,653 | |
| 1999 | 70 | 70 | ||
| Total | 1,905 | 1,985 | 2,516 | 6,406 |
The distribution of coho returns between the gear groups
has varied throughout the years (Table 5). The coho were harvested relatively
equally by trollers, gillnetters, and seiners in 1996 and 1998. Trollers
dominated the harvest of Old Franks coho in 1997. Only trollers and gillnetters
caught tagged coho in 1999.
Table 5. Old Franks coho bioenhancement contribution (number
of fish) to commercial fisheries by gear type as estimated by coded wire
tag recoveries.
| Gear Type | 1996 | 1997 | 1998 | 1999 | Total |
| Troll | 645 | 1,210 | 841 | 36 | 2,731 |
| Drift Gillnet | 699 | 375 | 767 | 34 | 1,875 |
| Purse Seine | 533 | 222 | 980 | 1,734 | |
| Sport | 65 | 65 | |||
| Total | 1,877 | 1,807 | 2,653 | 70 | 6,407 |
The commercial fisheries captured 29% of the Old Franks
coho in the northern part of Clarence Strait (district 106) when contribution
for all years is combined (Table 6). High numbers were also captured on
the outside of Baranof Island (district 113), the southeast side of Prince
of Wales Island (district 102) and the outside waters west of Prince of
Wales (district 104).
Table 6. Coho bioenhancement contribution (number of fish)
to fisheries by area harvested (ADF&G
statistical areas) and percentage of total harvest from all years by
area.
| Harvest Area (District) | 1996 | 1997 | 1998 | 1999 | Total | Overall % |
| Ketchikan Area (101) | 129 | 32 | 303 | 6 |
533
|
8.3
|
| southeast Prince of Wales (102) | 278 | 84 | 399 |
936
|
14.6
|
|
| west Prince of Wales, inside waters (103) | 40 | 36 | 55 | 8 |
139
|
2.2
|
| west Prince of Wales, outside (104) | 257 | 219 | 154 |
625
|
9.7
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|
| Sumner Strait (105) | 103 | 61 | 199 |
360
|
5.6
|
|
| northern Clarence Strait (106) | 693 | 374 | 730 | 34 |
1,834
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28.6
|
| Ernest Sound (107) | 14 |
14
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0.2
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|||
| Wrangell (108) | 2 |
2
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0.0
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|||
| Chatham Strait / Frederick Sound (109) | 79 | 155 | 118 |
355
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5.5
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|
| outside Baranof (113) | 168 | 461 | 296 | 13 |
898
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14.0
|
| Cross Sound / Icy Strait (114) | 27 | 12 |
39
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0.6
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||
| C. Spencer to C. Fairweather (116) | 9 | 23 |
32
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0.5
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||
| offshore Column Pt. to C. Ommaney (154) | 9 | 50 | 77 |
152
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2.4
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| area not reported | 132 | 268 | 104 | 8 |
488
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7.6
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| Total | 1,877 | 1,807 | 2,653 | 70 |
6,407
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100
|
Juvenile Coho in Neff Creek and New Pipe Creek
We marked reaches of Neff Creek and New Pipe Creek to be used as coho density monitoring sites in the upper watershed. Reaches of FP3 channel type short enough to be sampled with a depletion population estimate in one day were selected. Coho populations and densities in these two streams were lower than densities in Trail Creek and Toque Creek (Figure 6). Trail and Toque Creek were not sampled in 1999 but densities were likely similar or higher than in 1997 or 1998, consistent with the rest of the watershed. Coho densities are highest and probably at carrying capacities low in the Old Franks system, in the area of Trail Creek and possibly Toque Creek. Judging by coho densities, colonization is still occurring above the lakes. Neff Creek and New Pipe Creek both flow into upper Old Franks Creek on the downstream left bank at 1,675 and 3,564 meters up from the lake respectively.
Figure 6. Juvenile coho salmon densities in Old Franks
tributary streams.
Table 7 shows the summary statistics for Neff and New Pipe Creek for comparison with riparian habitat management objectives (RHMO’s).
Table 7. Stream survey summary statistics for Neff and
New Pipe Creeks, both FP3 channel type.
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| Neff Creek |
153.7
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8
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62
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5.4
|
121
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33
|
150
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0
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52
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0.3
|
3.6
|
9
|
20
|
38
|
| New Pipe Creek |
152.6
|
8
|
46
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5.9
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75
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19
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564
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33
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52
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0.3
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3.2
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13
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42
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94
|
Salmonid Abundance and Distribution in Upper Old Franks Creek
The 1999 juvenile snorkel survey included 49 habitat units between Upper Old Franks Lake and the top of anadromous habitat on Old Franks Creek. Coho densities were .11 fish per square meter (or 8.7 square meters of habitat per fish) in slow water (pool) habitats and .015 fish per square meter (or 65 square meters per fish) in fast water (riffle) habitats. These densities were the highest we have seen since the fishpass was installed (Figure 7). They were even higher than in 1995 when coho fry were planted in upper Old Franks Creek. Although 1999 had the highest coho densities we have observed, coho densities are probably still below carrying capacity in the creek. Densities should increase further in the next generation. Coho densities have been consistently over two times higher in slow water habitats than in fast water habitats.
We combined cutthroat and rainbow counts during snorkel surveys because the juveniles are nearly indistinguishable during snorkel surveys. The 1999 cuthroat/rainbow density was higher than all years since 1994 when the highest density was recorded. The 1994 density was still over twice the density recorded in 1999. Surveys in all years except 1997 were conducted between mid-July and mid-September, following cutthroat and rainbow fry emergence. The survey in 1997 was conducted in mid-June, prior to cutthroat and rainbow emergence so consisted entirely of age 1+ fish (Table 8). The density of age 1+ cutthroat/rainbow in 1997 was also the lowest since monitoring began.
The Dolly Varden density in 1999 was similar to 1998 when the highest density of all years was recorded. Dolly varden are difficult to find during snorkel surveys so they typically are significantly under counted. They seem to be more active and visible during high water when other species are harder to count and snorkeling conditions are more difficult.
Figure 7. Fish densities in Upper Old Franks Creek.
Table 8. Percentage of age 1+ (versus age 0) cutthroat/rainbow
(>50 mm) and coho (>55 mm) counted during snorkel surveys.
| Aug/1994 | Aug/1995 | Sep/1996 | Jun/1997* | Jul/1998 | Aug/1999 | |
| Cut/Rnb | 23% | 25% | 30% | 100% | 42% | 64% |
| Coho | 0% | 13% | 12% | 19% | 12% |
* The 1997 survey was conducted in mid-June, prior to cutthroat and rainbow emergence.
Based on qualitative observations, coho appeared to be more associated with some sort of cover (undercut banks, rootwads, tree branches) with lower water velocities than were the cutthroat/rainbow. The small cutthroat/rainbow (50-150mm) were often in faster water currents, but where there were large congregations of juvenile coho (>20) there were usually a few cutthroat/rainbow mixed in. The large cutthroat/rainbow (> 150mm) were generally in the largest, deepest pools with or without cover; the same areas that the adult coho congregated. Newly emergent fry were most abundant along the stream margins in very shallow (<10cm) water.
Tier 3 Stream Survey of Upper Old Franks Creek
We conducted the tier 3 survey to mark new habitat units for fish density measurements. The survey determined total usable habitat area in the stream and segregated out specific habitat types. The total usable area at a baseflow condition was 113,238 square meters (Table 9). The split between slow water and fast water area showed a lower proportion of slow water habitat in 1999 than in past surveys. This is because the new survey protocol classifies pools based on specifically defined hydraulic characteristics (residual depth and minimum area) so some areas that may have appeared to be pools in the past actually key out as riffles. The total area surveyed was higher in 1994 and 1999 than in prior years because these surveys were more thorough and covered all stream area to the top of anadromous habitat.
Table 9. Comparison of total macro habitat unit areas
in Upper Old Franks Creek by year.
| Year | Slow Water | Fast Water | Total Area | % slow water |
| 1990 | 55,708 | 39,140 | 94,848 |
59
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| 1991 | 46,088 | 42,178 | 89,658 |
51
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| 1992 | 49,243 | 45,679 | 94,922 |
52
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| 1994 | 58,435 | 54,184 | 112,989 |
52
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| 1999 | 49,379 | 63,502 | 113,238 |
44
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Table 10 shows summary elements from the habitat survey
data. These data elements will be used to compare to regional riparian
habitat management objectives (RHMO’s) and can be an indicator of stream
and fish habitat condition. The RHMO’s have not yet been derived for use
with the latest protocol.
Table 10. Upper Old Franks Creek stream survey summary
statistics, 1999.
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| FP5 |
138.5
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0.0
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1266.1
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11
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111
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29
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838
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62
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212
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21
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9
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0.3
|
4.0
|
26
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| LC1 |
0.0
|
1266.1
|
375.8
|
8
|
67
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21
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206
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28
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72
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11
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21
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0.4
|
2.3
|
31
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| FP5 |
402.5
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1641.9
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2570.7
|
35
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100
|
24
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1404
|
50
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258
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26
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14
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0.3
|
3.0
|
24
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| MC2 |
0.0
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4173.5
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612.3
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20
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67
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19
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374
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30
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73
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5
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33
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0.6
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1.6
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28
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| FP5 |
84.0
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4800.6
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557.4
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10
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88
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19
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274
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47
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205
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34
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18
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0.3
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2.9
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22
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| FP4 |
533.7
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5358.0
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1669.6
|
37
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64
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18
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842
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33
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242
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69
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22
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0.4
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2.6
|
28
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| MC2 |
0.0
|
7027.6
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455.6
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12
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79
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12
|
108
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25
|
86
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22
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26
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0.3
|
3.2
|
15
|
Discussion
The coho escapement in 1999 was the highest since the fishpasses were installed. The 1999 escapement was significantly higher than only the 1993 – 1995 escapements. The 95% confidence intervals are wide because we had a small number of recaptures relative to the number of marked coho. Coho production and escapement during this initial colonization period have exceeded expectations for the first generation of returns. Escapements should continue to increase as not all rearing habitat in the upper watershed is yet fully seeded.
The 1999 return included mostly naturally spawned Old Franks coho. No coded wire tagged coho were captured or observed during escapement monitoring activities. The 1999 return included offspring from coho produced by the first year of bioenhancement (brood year was 1996). Most production returning in 1999 can be attributable to coho that used the fishpasses.
Coho salmon juveniles are present throughout the accessible habitat in the Old Franks watershed, although all habitat is not yet fully seeded. The few small tributary streams, for example Toque Creek and Trail Creek, flowing into the lakes and lower Old Franks Creek appear fully seeded. Toque and Trail Creek densities are in the range of .4 – 1.0 coho per square meter in September. The snorkel surveys in upper Old Franks Creek indicate that habitats in that area currently have coho densities of 1/2 to 1/8 that of similar stream channels on southern Prince of Wales Island (Craig District unpublished data). Rearing densities in the lakes are unknown but minnow trapping and fyke netting catch rates indicate that densities are high. It appears, judging by observed rearing densities, that the upper Old Franks Creek area (above the lakes) has the capability to produce more coho than are now present.
Sports fishing occurred at the mouth of Old Franks Creek in 1999, mostly by local residents in the Skowl Arm area. Fishing activity seemed lower than in 1997 when anglers were observed fishing at the mouth on many occasions.
Recreational use of the upper Old Franks area has increased since the logging roads were built into the watershed. Canoeists brushed a trail from the end of the road to a point on Old Franks Creek to allow easier access for hauling canoes between the road and the creek. Fishing opportunities in the upper watershed are primarily limited to resident fish. Most coho turn relatively dark by the time they reach the upper watershed in late September and October so are less desirable than fresh fish available near the mouth.
Logging activities in the upper watershed continued in
1999. Approximately 2.5 miles of new road were completed. Harvest operations
were initiated on 14 new logging units. These activities should be completed
in 2000. Access to the upper watershed will probably be limited again in
2000 by the logging activities.
Conclusions
Acknowledgements
This work could not have been completed without the faithful field
work and dedication of Susan Farzan, Chad Fealko, and Tim Paul.
Contents of past Old Franks reports
The following is a list of related work done in the Old Franks Watershed and the years covered:
1978-1989 - Zadina and Haddix 1992. Summary of work from 1978 - 1989, lake physical characteristics, water chemistry, and bathymetric measurements, zooplankton sampling, resident fish sampling, anadromous species production potential estimates, bioenhancement recommendations
1990 - Bryant 1991. Upper Old Franks Creek habitat and fish survey, lake zooplankton densities, resident fish surveys in Old Franks and Upper Old Franks Lakes, discussion of possible species interactions following anadromous introductions
1994 - Hannon. 1995. First bioenhancement coho release, escapement estimate, watershed analysis summary, stream survey summary of upper Old Franks Creek, upper Old Franks Creek fish survey
1994 - USFS 1994. Watershed Analysis - physical and biological characteristics of the watershed upstream of Upper Old Franks Lakes, riparian habitat conservation areas identified, effects of the logging on fish habitat evaluated
1995 - Hannon. 1996. Escapement data, resident fish surveys in Lake Mary and Old Franks Lake, upper Old Franks Creek fish survey, electrical fish counter accuracy test
1996 - Hannon. 1997. First year of bioenhancement returns, coded wire tag results, escapement data, resident fish surveys in Lake Mary and upper Old Franks Lake, stream surveys of Trail Creek and Toque Creek, upper Old Franks fish survey
1993-1996 - Denton 1997. Summary of the coho bioenhancement work completed, egg take methods and results, incubation and rearing, tagging and release data, transport and planting, coded wire tag results from 1996
1997 - Hannon. 1997b. Escapement data, coded wire tag results, Trail Creek and Toque Creek coho population data, upper Old Franks Creek fish survey
1998 – Hannon. 1998. Escapement data, coded wire
tag results, Trail Creek and Toque Creek coho population data, upper Old
Franks Creek fish distributions
Literature Cited
Coghill, K. 1997. Draft fish habitat and stream survey procedures for the Tongass National Forest. June 20, 1997 version. FSL Juneau. 50p.
Hannon, J. 1998. Old Franks Monitoring--1998. Craig Ranger District. USFS Craig. 26p.
Denton, C. 1997. Final report Old Franks Lakes coho bioenhancement project. Alaska Department of Fish and Game CFMD Division. Ketchikan. 20p.
Thedinga, J.F. and S.W. Johnson. 1995. Retention of jet-injected marks on juvenile coho and sockeye salmon. Transactions of the American Fisheries Society 124:782-785.
USDA Forest Service. 1999. Aquatic ecosystem management handbook, chapter 20 - fish and aquatic stream habitat survey. USFS Juneau. 50p.
USDA Forest Service. 1994. Watershed analysis for Old Franks Creek Watershed. Ketchikan Area, Ketchikan, AK. 70p.
White, G., D. Anderson, K. Burnham, D. Otis. 1982. Capture-recapture and removal methods for sampling closed populations. LA-8787-NERP. Los Alamos National Laboratory. Los Alamos, NM. 235p.
Zadina and Haddix. 1992. Fisheries habitat evaluation and limnological investigations of the Old Franks Lake system, Prince of Wales Island, SE Alaska 1978 - 1989. ADF&G FRED Division. Juneau, AK. 37p.
Old Franks Data Files
Data collected by the Craig Ranger District in 1999 resides on the Craig server at /unit/fw/fish/Oldfranks/1999. Individual files are:
Data files for years prior to 1999 are in the following folders /unit/fw/fish/Oldfranks/1998/, /1997/, /1996/, /1995/, /1994/, and /pre 1994/.